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FIG. 66.-Diagram of the morphological relations of the Astacina.

form which characterize each of the genera and specics, would appear in the place of the names of the former, or of the circles which represent the latter. All these figures would represent abstractions - mental images which have no existence outside the mind. Actual facts would begin with drawings of individual animals, which we may suppose to occupy the place of the dots above the upper line in the diagram.

That all crayfishes may be regarded as modifications of the common plan A, is not an hypothesis, but a generalization obtained by comparing together the observations made upon the structure of individual crayfishes. It is simply a graphic method of representing the facts which are commonly stated in the form of a definition of the tribe of crayfishes, or Astacina.

This definition runs as follows:

Multicellular animals provided with an alimentary canal and with a chitinous cuticular exoskeleton; with a ganglionated central nervous system traversed by the œsophagus; possessing a heart and branchial respiratory organs.

The body is bilaterally symmetrical, and consists of twenty metameres (or somites and their appendages), of which six are associated into a head, eight into a thorax, and six into an abdomen. A telson is attached to the last abdominal somite.

The somites of the abdominal region are all free, those of the head and thorax, except the hindermost, which is

DEFINITION OF THE GROUP ASTACINA,

255

partially free, are united into a cephalothorax, the tergal wall of which has the form of a continuous carapace. The carapace is produced in front into a rostrum, at the sides into branchiostegites.

The eyes are placed at the ends of movable stalks. The antennules are terminated by two filaments. The exopodite of the antenna has the form of a mobile scale. The mandible has a palp. The first and second maxillæ are foliaceous; the second being provided with a large scaphognathite. There are three pairs of maxillipedes, and the endopodites of the third pair are narrow and elongated. The next pair of thoracic appendages is much larger than the rest, and is chelate, as are the two following pairs, which are slender ambulatory limbs. The hindmost two pairs of thoracic appendages are ambulatory limbs, like the foregoing, but not chelate. The abdominal appendages are small swimmerets, except the sixth pair, which are very large, and have the exopodite divided by a transverse joint.

All the crayfishes have a complex gastric armature. The seven anterior thoracic limbs are provided with podobranchiæ, but the first of these is always more or less completely reduced to an epipodite. More or fewer arthrobranchiæ always exist. Pleurobranchiæ may be

present or absent.

In this tribe of Astacina there are two families, the Potamobiida and the Parastacide; and the definition of each of these families is formed by superadding to the

definition of the tribe the statement of the special pecu. liarities of the family.

Thus, the Potamobiidæ are those Astacina in which the podobranchia of the second, fourth, fifth, and sixth thoracic appendages are always provided with a plaited lamina, and that of the first is an epipodite devoid of branchial filaments. The first abdominal somite invariably bears appendages in the males, and usually in both sexes. In the males these appendages are styliform, and those of the second somite are always peculiarly modified. The appendages of the four following somites are relatively small. The telson is very generally divided by a transverse incomplete hinge. None of the branchial filaments are terminated by hooks; nor are any of the coxopoditic setæ, or the longer setæ of the podobranchiæ hooked, though hooked tubercles occur on the stem and on the lamina of the latter. The coxopoditic setæ are always long and tortuous.

In the Parastacidæ, on the other hand, the podobranchiæ are devoid of more than a rudiment of a lamina, though the stem may be alate. The podobranchia of the first maxillipede has the form of an epipodite; but, in almost all cases, it bears a certain number of well developed branchial filaments. The first abdominal somite possesses no appendages in either sex: and the appendages of the four following somites are large. The telson is never divided by a transverse hinge. More or fewer of the branchial filaments of the podo

ALLIES OF THE CRAYFISH.

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branchiæ are terminated by short hooked spines; and the coxopoditic setæ, as well as those which beset the stems of the podobranchiæ, have hooked apices.

The definitions of the genera would in like manner be given by adding the distinctive characters of each genus to the definitions of the family; and those of the species by adding its character to those of the genus. But at present it is unnecessary to pursue this topic further.

There are no other inhabitants of the fresh waters, or of the land, which could be mistaken for crayfishes; but certain marine animals, familiar to every one, are so strikingly similar to them, that one of these was formerly included in the same genus, Astacus; while another is very often known as the "Sea-crayfish." These are the "Common Lobster," the "Norway Lobster," and the "Rock Lobster or" Spiny Lobster."

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The common lobster (Homarus vulgaris, fig. 67) presents the following distinctive characters. The last thoracic somite is firmly adherent to the rest; the exopodite of the antenna is so small as to appear like a mere movable scale; all the abdominal appendages are well developed in both sexes; and, in the males, the two anterior pairs are somewhat like those of the male Astacus, but less modified.

The principal difference from the Astacina is exhibited by the gills, of which there are twenty on each side; namely, six podobranchiæ, ten arthrobranchiæ, and four

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